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Ephemeroptera: Heptageniidae of Gunnison County, Colorado

Rhithrogena undulata
Small Western Red Quill, Red Quill

(Banks) 1924

Updated 6 Feb 2012
TSN 100591

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On this website:
Introduction to Rhithrogena

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Mayflies of Colorado distribution map (USGS) Coordinated by Boris Kondratieff


Banks, N. 1924 Descriptions of new neuropteroid insects. Bulletin of the Museum of Comparative Zoology. 65(12):419-455, pl. 1-4.
     Described as Epeorus undulatus on page 425, with an illustration (Fig 57) of the male genitalia (ventral view):

Edmunds GF Jr. 1952b Studies on the Ephemeroptera. Part II. The taxonomy and biology of the mayflies of Utah Unpublished Ph.D. Thesis, University of Massachusetts 399 pages.

Edmunds GF Jr. 1995 Habitat differences between northern and southern populations of mayflies of the western United States. Pages 171-176 in Corkum LD; Ciborowski JJH. Current Directions in Research on Ephemeroptera. Canadian Scholars' Press, Inc. Toronto.
     Copied from pages 172 and 173:

Jensen,SL 1966 The Mayflies of Idaho (Ephemeroptera). M.S. Thesis, University of Utah, Utah. 364 p.
     Quote from page 197: "Banks (1924) described this species from a male imago collected in New Mexico. Traver (1935b) and Edmunds(1952b) provide descriptions of the adults. Of the nymphs, Edmunds (1952b:178) states concerning the description:
{Posterior margin of head moderately concave, otherwise identical with R. doddsi [R. hageni Eaton]}
Taxonomy and Biology: According to Traver (1935b), R. hageni [probably a typo, Jensen intended to say R. undulata] is a member of the jennuna-undulata species group, distinquished by lacking basal lateral spines on the penes of the male. The nymphs, however, are extremely difficult to distinguish from R. hageni and R. morrisoni (Banks).
Nymphs of this species are usually collected in large, silted streams at elevations of from 4,000 to 6,000 feet, where they live on rocks and debris in the slower water. "

McCafferty,WP; Durfee,RS; Kondratieff,BC 1993 Colorado mayflies (Ephemeroptera): an annotated inventory. Southwestern Naturalist 38 3, 252-274. PDF
     Quote from page 262: "This species appears to be limited to the western slope in Colorado, and it shows considerable color variation throughout its range. It, as well as R. plana and R. vitta described by Allen and Chao (1987a) from Arizona, will likely prove to be variants of R. morrisoni (Banks). Rhithrogena morrisoni has one page priority over R. undulata (see Banks, 1924) and has been reported from Arizona, California, Nevada, New Mexico, and Utah. In addition, the first author has seen specimens from Idaho and Wyoming. "

McCafferty,WP and Provonsha, AV The Mayflies of North AmericaSpecies List (Version 8Feb2011)
     Here is the geographic range and synonyms:
Rhithrogena undulata (Banks), 1924 [CAN:FN,NE,NW;USA:NE,NW,SW]
    * Epeorus undulata Banks, 1924 (spell.)
    * Epeorus undulatus Banks, 1924 (orig.)

Brown,WS 2004 Mayflies (Ephemeroptera) of Gunnison County, Colorado, USA