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Trichoptera of Gunnison County, Colorado

Introduction to the caddis family Ueonidae
Uenoid Case Makers

Iwata, 1927
Updated 16 Feb 2019
TSN 568757

Three living Neothremma alicia larvae are crawling on moss under a thin film of water. They have cases of fine stones that are blurred by the fast cold water.

The three animals on the right were photographed in a small subalpine stream on 18 July 2007.
They have fine stone cases, which are obscured by the water and reflection.
In this stream, Neothremma alicia crawl around and burrow head first into the fronds of moss.

Provisional Species List

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Family Overview - University of Alberta Entomology Collection Family page
     Has habitat, identification, life history, conservation and more.


Herrmann,SJ; Ruiter,DE and Unzicker,JD 1986 Distribution and records of Colorado Trichoptera. Southwestern Naturalist 31(4), 421-457.

Iwata,M 1927 Trichopterous larvae from Japan. Annotationes Zoologicae Japonenses, 11, 203-233.
     The author described the family Ueonidae on page 214 for the first time.

Ross, H. H. 1949. The caddisfly genus Neothremma Banks (Trichoptera: Limnephilidae). Journal of the Washington Academy of Sciences 39:92-93.

Vineyard,RN and Wiggins,GB 1988 Further revision of the caddisfly family Uenoidae (Trichoptera): evidence for inclusion of Neophylacinae and Thremmatidae. Systematic Entomology 13(3) 361-372.
     Abstract: "A survey of characters within the Limnephilidae was made to test the cladistic hypothesis underlying traditional classification placing Neophylacinae as a subfamily of the Limnephilidae. Characters emerged which support an alternative hypothesis of phylogeny and classification, linking the Neophylacinae with the Thremmatidae and with the Uenoidae. The Uenoidae as defined here contain two monophyletic groups: subfamily Uenoinae containing the four genera Uenoa, Sericostriata, Neothremma and Farula previously constituting Uenoidae (sensu Wiggins et al., 1985), and subfamily Thremmatinae consisting of Thremma, Neophylax and Oligophlebodes. The new classification is shown to have predictive value for behaviour and ecology of the taxa."

Wiggins,GB 1996 Larvae of the North American Caddisfly Genera (Trichoptera). 2nd Edition. University of Toronto Press, 457 pages.

Wiggins,GB and Erman,NA 1987 Additions to the systematics and biology of the caddisfly family Uenoidae (Trichoptera). Canadian Entomologist 119:867-872.
     Abstract: " A description is given for the male, female, and larva of Farula praelonga sp. nov. and for the female and larva of Neothremma genella Denning. Farula praelonga sp. nov. is distinctive in having an exceptionally long internal branch of segment X. The female of Neothremma genella is the only female yet known in that genus to have greatly reduced ventrolateral lobes of sternum IX. Modifications are made to the keys previously published for adults of Farula and Neothremma to incorporate this new information. These two species occur together in abundance; they are univoltine and have emergence periods 6 months apart."

Wiggins,GB; Weaver,JS; Unzicker,JD 1985 Revison of the caddisfly family Uenoidae (Trichoptera). Canadian Entomologist 117, 763-800.
     Abstract: "The western Nearctic genera Neothremma and Farula are assigned to the family Uenoidae, previously comprising only the type genus Uenoa from Asia; a new genus Sericostriata with S. surdickae n. sp. from western North America is also added to the family. Traditionally assigned to the Limnephilidae where they have been always anomalous, Neothremma and Farula are shown to share many synapomorphic larval characters with Uenoa; under close examination, characters of the adults support this relationship. Familial and generic diagnoses are provided, reflecting the revised composition of the Uenoidae. Generic keys to adults, larvae, and pupae are given; species are briefly reviewed and keys given to adults; Uenoa arcuata n. sp. is described from Assam, Uenoa janetscheki Botosaneanu is placed as a junior subjective synonym under U. hiberna Kimmins, and Neothremma laloukesi Schmid as a junior subjective synonym of N. alicia Dodds and Hisaw. Phylogenetic relationships among the genera are considered. This study is an instructive example of the importance of larvae in providing data for the systematics of Trichoptera."

Wiggins,GB and Wisseman,RW 1992. New North American species in the genera Neothremma and Farula, with hypotheses on phylogeny and biogeography (Trichoptera: Uenoidae). Canadian Entomologist 124:1063-1074.
     Abstract: " Three new species are described in the caddisfly family Uenoidae: Neothremma prolata, from Hood River County, Oregon; Neothremma mucronata from Lassen County, California; and Farula constricta from Multnomah County, Oregon. Following examination of the holotypes of several species, misinterpretation of the male genitalia morphology of Farula wigginsi Denning is corrected, leading to the recognition of that name as a junior synonym of F. petersoni Denning. Interpretation of male genitalic morphology in the original description of F. geyseri Denning is revised. Phylogenetic relationships are inferred from male genitalic morphology for the species of Neothremma and Farula. Biogeographic patterns of the species in both genera are highly congruent with the phylogenies."

Brown,WS 2005 Trichoptera of Gunnison County, Colorado, USA