Updated 29 Feb 2016
Good LinksOn this website:
Introduction to Rhyacophila
Photos, Map, Museum specimens, DNA - Barcodinglife.org
University of Alberta Entomology Collection Species page
Has illustration of male genitalia, description, habitat information, range and more.
Locations CollectedIllinois Natural History Survey (INHS) Database shows this species present in North Quartz Creek. Allan (1975) reports this species from Cement Creek.
ReferencesAllan,JD 1975 The distributional ecology and diversity of benthic insects in Cement Creek, Colorado. Ecology 56:1040-1053. PDF
Erman,NA 1989 Species composition, emergence, and habitat preferences of Trichoptera of the Sagehen Creek Basin, California, USA. The Great Basin Naturalist, 186-197. PDF
Quote from page 188: "Four other species (Desmona bethula, Rhyacophila ardala, R. vaccua, and R. verrula) were occasionally trapped in November and December." The author found Rhyacophila verrula in springs and spring-fed streams.
Giersch, JJ 2002 Revision and phylogenetic anaysis of the verrula and alberta species groups of Rhyacophila Pictet 1834 with description of a new species (Trichoptera: Rhyacophilidae). Master of Science Thesis. Montana State University http://virgin.msu.montana.edu/Joe/
Herrmann,SJ; Ruiter,DE and Unzicker,JD 1986 Distribution and records of Colorado Trichoptera. Southwestern Naturalist 31 4, 421-457.
They note the habitat for this species is streams and rivers, the altitudinal range is 1707 to 3048m and adult collection dates are 13 July to 19 October. Quote from page 452: "Smith (1968) reported this species from Colorado but gave no collection data. He also pointed out that, in Idaho, R. verrula was always found in association with a green algae, Prasiola sp., which it ate." They list this species as present in Gunnison county.
Milne,LJ 1936 Studies in North American Trichoptera. Part 3. Cambridge, Mass. Author's publication. 128 pages
Nimmo, A 1971 The adult Rhyacophilidae and Limnephilidae (Trichoptera) of Alberta and eastern British Columbia and their post glacial origin. Quaestiones Entomologicae 73: 3-234. Sometimes you can buy a used copy at Amazon.com
Peckarsky,BL 1980 Influence of detritus on colonization of stream invertebrates. Canadian Journal of Fisheries and Aquatic Sciences 37, 957-963.
Schmid,F 1970 Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada 66:1-230.
Short,RA and Ward,JV 1980 Macroinvertebrates of a Colorado high mountain stream. The Southwestern Naturalist, 23-32. PDF
Smith,SD 1968 The Rhyacophila of the Salmon river drainage of Idaho with special reference to larvae. Annals of the Entomological Society of America 61 3, 655-674.
Has a key to larvae that includes R. verrula. Also contains illustrations of R. verrula's head, pronotum, anal proleg and mandibles (Figure 13). Peak emergence of adults was during September and October in the Salmon River drainage of Idaho between 4000 and 8000 feet. R. verrula collected from a spring site had larvae of all ages present at the same time, probably due to the constant conditions of the spring. Smith always found larvae associated with the green algae Prasiola sp. Gut analysis showed R. verrula larvae ate algae, watercress and unidentifiable plant material. This is unique among North American Rhyacophila where most species are carnivorous, at least in the later instars.
Brown, WS 2004 Trichoptera of Gunnison County, Colorado, USA