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Stoneflies - Plecoptera: Perlodidae

Isogenoides colubrinus (Hagen 1874)

Blackfoot Springfly
Updated 5 August 2017


Older publications may refer to this species as Isogenus colubrinus.

Good Links

On this website:
Introduction to Isogenoides
Key to Perlodidae Nymphs

Other Websites:
Photos, Map, Museum specimens, DNA - Barcodinglife.org


Baumann,RW, Gaufin,AR and Surdick,RF 1977 The stoneflies (Plecoptera) of the Rocky Mountains. Memoirs of the American Entomological Society 31, 1-208. PDF
     Quote from page 127: "This species inhabits large rivers. The adults emerge from March to August. "

Barton,DR 1980 Observations on the life histories and biology of Ephemeroptera and Plecoptera in northeastern Alberta. Aquatic Insects, 2(2) 97-111. PDF

Hagen,HA, 1874 Report on the Pseudo-neuroptera and Neuroptera collected by Lieut. W.L. Carpenter in 1873 in Colorado. Annual Report of the U.S. Geological and Geographical Survey of the Territories, embracing Colorado, 7: 571-577.
     Described as Isogenus colubrinus.

Hanson,JF 1943 Records and Descriptions of North American Plecoptera. Part III. Notes on Isogenoides. American Midland Naturalist, 29(3) 657-669.

Klapálek, Frantisek 1912 Plécoptères. I. Fam. Perlodidae; [monographische Revision. II. Fam. Perlidae; Subfam. Perlinae, Subfam. Neoperlinae; mongraphische Revision] Series Sélys-Longchamps, Edmond de, baron, 1813-1900. Collections zoologiques; catalogue systematique et descriptif, fasc. 4, pt. 1-2.
     Discussed as Isogenoides colubrinus.

Kondratieff,BC and Baumann,RW 2002 A review of the stoneflies of Colorado with description of a new species of Capnia (Plecoptera: Capniidae). Transactions of American Entomological Society 128 3, 385-401.
     Quote from page 397: "This species, although not as commonly collected as the following species [Isogenoides elongatus and Isogenoides zionensis], occurs in similar larger rivers of the plateau region. "

Sandberg,JB and Stewart,KW 2004 Capacity for extended egg diapause in six Isogenoides Klapálek species (Plecoptera: Perlodidae). Transactions of the American Entomological Society 130(4): 411-423.
     Abstract: "The eggs of six Isogenoides species have been incubated at a single, approximately simulated, San Miguel River, Colorado, seasonal temperature regime for 2-5 years. Eggs were collected from reared, laboratory-mated females of I. colubrinus, I. doratus, I. elongatus, I. frontalis, I. varians, and I. zionensis. Eggs were held in 8 ml containers and visually examined usually weekly for development and hatch. Only some I. zionensis eggs from a combined first and second mass of a Leopard Creek, Colorado population hatched directly within two weeks, continuing until late September, and resuming mainly May-June the following year; one and two eggs hatched in 2002 and 2004, respectively. I. doratus and I. varians experienced hatch only after an over-summer, 3-4 month diapause, and I. varians again in late August-September during the second year. Populations of I. colubrinus, I. elongatus, I. frontalis and a San Miguel River, Colorado population of I. zionensis began hatching only in the year following deposition, after a 10-12 month diapause. Some hatching of these four species occurred at spring-summer temperatures in subsequent years (2nd, 3rd, 3rd, and 4th years, respectively), with no intervening hatch at winter temperatures. The study confirms extended, usually asynchronous diapause and hatching for from 3 months to 4 years, probably genetically controlled, in the six species, and a great degree of adaptive capacity for diapause in the genus Isogenoides."

Sandberg,JB and Stewart,KW 2005a Vibrational communication (Drumming) of the nearctic stonefly genus Isogenoides (Plecoptera: Perlodidae). Transactions of American Entomological Society 131 (1+2) 111-130. PDF
     Has sonogram of drumming duet between male and female on page 125.

Sandberg,JB and Stewart,KW 2005b Holomorphology and systematics of the stonefly genus Isogenoides (Plecoptera: Perlodidae). Transactions of American Entomological Society 131 (3+4) 269-345.


subgenital plate

Brown, WS 2004 Stoneflies or Plecoptera of Gunnison County, Colorado, USA