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Ephemeroptera: Ephemerellidae of Gunnison County, Colorado

Introduction to the Mayfly genus Ephemerella
Pale Morning Dun, PMDs, Pale Morning Olive, Pale Morning Spinner, Pale Olive Quill

Walsh 1862
Updated 19 January 2019
TSN 101233

Provisional Species List

Ephemerella aurivillii
Ephemerella excrucians (inermis)
Ephemerella dorothea infrequens


This genus has many lovely translucent little yellow species. The taxonomy has been changing recently, but no matter, the fishermen recognize the local species of Pale Morning Dun in streams across our planet.

Good Links

On this website:
Ephemerella apopsis
Ephemerella inermis
Introduction to Ephemerellidae

Other Websites:
Photo Adult male - go to the bottom of the page and among many great photos, there is a Pale Morning Dun or Ephemerella dorothea second from the right.


Allen,RK 1968 New species and records of Ephemerella (Ephemerella) in western North America (Ephemeroptera: Ephemerellidae). Journal of the Kansas Entomological Society 41(4):557-567 PDF

Brinkman,SF and Johnston,WD 2012 Acute toxicity of zinc to several aquatic species native to the Rocky Mountains. Archives of environmental contamination and toxicology, 62(2), 272-281.

Gill,BA; Harrington,RA; Kondratieff,BC; Zamudio,KR; Poff,NL and Funk,WC 2014 Morphological taxonomy, DNA barcoding, and species diversity in southern Rocky Mountain headwater streams. Freshwater Science 33(1) 288-301.
     Working in wadeable streams on the Front Range of Colorado, they found a cryptic species of Ephemerella alongside E. dorothea and infrequens.

Jacobus,LM and McCafferty,WP 2003 Revisionary contributions to North American Ephemerella and Serratella (Ephemeroptera: Ephemerellidae). Journal of the New York Entomological Society 111:174-193. PDF
     This paper reorganized the taxonomy map for the genus Ephemerella.

Jacobus, LM and McCafferty, WP 2008 Revision of Ephemerellidae genera (Ephemeroptera). Transactions of the American Entomological Society 134: 185-274. PDF

Short,RA; Canton,SP and Ward,JV 1980 Detrital processing and associated macroinvertebrates in a Colorado mountain stream. Ecology, 61(4), 727-732. PDF

Walley GS. 1930. Review of Ephemerella nymphs of western North America (Ephemeroptera). Canadian Entomologist 62(1):12-20, pl. 2-3. PDF

Walsh, BD 1862 List of the Pseudoneuroptera of Illinois contained in the cabinet of the writer, with descriptions of over forty new species, with notes on their structural affinities. Proceedings of the Academy of Natural Sciences Philadelphia 14:361-402.
     Walsh described the genus Ephemerella in this paper.
Walsh 1862 decription of the mayfly genus Ephemerella

"It ain't what you don't know that gets you into trouble.
It's what you know for sure that just ain't so."
--Mark Twain

Brown,WS 2009 Ephemeroptera of Gunnison County, Colorado